Atlantic Salmon Ecosystems Research Team Publications


Olmos M, Payne MR, Nevoux M, Prévost E, Chaput G, Du Pontavice H, Guitton J, Sheehan TF, Mills K, Rivot E. 2020. Spatial synchrony in the response of a long range migratory species (Salmo salar) to climate change in the North Atlantic Ocean. Global Change Biol. 26(3): 1319-1337.

Nieland JL, Sheehan TF. 2020. Quantifying the effects of dams on Atlantic salmon in the Penobscot River watershed, with a focus on Weldon Dam. 19-16; 90 p.


Bronger K, Sheehan TF. 2019. Workshop Report: Introduction and Overview of the ROAM (RAFOS Ocean Acoustic Monitoring) Approach to Marine Tracking . 19-15; 82 p.

Sheehan TF. 2019. Current status of knowledge, data, and research efforts on Atlantic salmon at Greenland: what do we have, what do we need, and what should we do moving forward?. 19-10; 89 p.

Hare JA, Kocik JF, Link JS. 2019. Atlantic salmon recovery informing and informed by ecosystem‐based fisheries management. Fisheries. 44(9): 403-411.

Hawkes JP, Goullete GS, Moctezuma AM, Atkinson EJ, Cox ON. 2019. Supplementation of Atlantic salmon in the southern extent of their range: Evaluation of age-1 hatchery smolt stocking in a small coastal watershed. In: Park J, Tessier L, editors. Second NPAFC-IYS Workshop on Salmon Ocean Ecology in a Changing Climate 15: 197–199.

Staudinger MD, Mills KE, Stamieszkin K, Record NR, Hudak CA, Allyn A, Diamond A, Friedland K, Golet W, Henderson ME, Hernandez CM, Huntington TG, Ji R, Johnson CL, Johnson DS, Jordaan A, Kocik J, Li Y, Liebman M, Nichols OC, Pendleton D, Richards RA, Robben T, Thomas AC, Walsh HJ, Yakola K. 2019. It’s about time: A synthesis of changing phenology in the Gulf of Maine ecosystem. Fish Oceanogr. 2019: 1–34.

Borggaard DL, Dick DM, Star J, Alexander M, Bernier M, Collins, M, Damon-Randall K, Dudley R, Griffis R, Hayes S, Johnson M, Kircheis D, Kocik J, Letcher B, Mantua, N, Morrison, W, Nislow K, Saba V, Saunders R, Sheehan T, Staudinger MD. 2019. Atlantic salmon (Salmo salar) climate scenario planning pilot report. 19-05; 89 p.

Lipsky CA, Saunders R, Stevens JR, O’Malley M, Music P. 2019. Developing sampling strategies to assess the Penobscot River estuary (2010-2013). 19-02; 56 p.

Stevens JR, Sheehan TF, Kocik JF. 2019. Modeling the impacts of dams and stocking practices on an endangered Atlantic salmon Salmo salar population in the Penobscot River, Maine, USA. Can J Fish Aquat Sci. 76(10): 1795-1807.

Vendramin N, Cuenca A, Sørensen J, Alencar ALF, Christiansen DH, Jacobsen JA, Axen C, Lieffrig F, Ruane NM, Martin P, Sheehan T, Iburg TM, Rimstad E, Olesen NJ. 2019. Presence and genetic variability of Piscine orthoreovirus genotype 1 (PRV‐1) in wild salmonids in Northern Europe and North Atlantic Ocean. J Fish Dis. 42(8): 1107-1118.

Renkawitz MD, Saunders R, Bailey SL. 2019. Program and abstracts from the 2018 Atlantic Salmon Ecosystems Forum. 19-09; 84 p.

Sheehan TF, Barber J, Dean A, Deschamps D, Gargan P, Goulette G, Maxwell H, Tavner I, Nygaard R, Bradbury IR, Robertson MJ, Ó Maoiléidigh N. 2019. The International sampling program: Continent of origin and biological characteristics of Atlantic salmon collected at West Greenland in 2017. 19-12; 44 p.

Arnekleiv JV, Davidsen JG, Sheehan TF, Lehnert SJ, Bradbury IR, Rønning L, Sjursen AD, Kjӕrstad G, Lubinski BA, Nilssen KJ. 2019. Demographic and genetic description of Greenland's only indigenous Atlantic salmon Salmo salar population. J Fish Biol. 94(1): 154-164.


Jeffery NW, Wringe BF, McBride MC, Hamilton LC, Stanley RRE, Bernatchez L, Clément M, Gilbey J, Sheehan TF, Bradbury IR. 2018. Range-wide regional assignment of Atlantic salmon (Salmo salar) using genome wide single-nucleotide polymorphisms. Fish Res. 206: 163-175.

Sheehan TF, Deschamps D, Downie H, Hawkes J, McAuliffe M, Millane M, Sims K, Nygaard R, Lubinski B, Robertson MJ, Maoiléidigh NO. 2018. The International Sampling Program: Continent of origin and biological characteristics of Atlantic salmon collected in West Greenland in 2016. 18-15; 29 p.

Liebich KB, Kocik JF, Taylor WW. 2018. Reclaiming a space for diadromous fish in the public psyche and sense of place. Fisheries. 43(5): 231-240.


Meseck SL, Li JY, Sunila I, Dixon M, Clark P, Music P, Lipsky C, Stevens JR, Wikfors GH. 2017. Characterizing seston in the Penobscot River Estuary. Mar Environ Res. 131: 43-56.

Dixon HJ, Dempson JB, Sheehan TF, Renkawitz MD, Power M. 2017. Assessing the diet of North American Atlantic salmon (Salmo salar L.) off the West Greenland coast using gut content and stable isotope analyses. Fish Oceanogr. 26(5): 555-568.

Hawkes JP, Sheehan TF, Stich DS. 2017. Assessment of early migration dynamics of river-specific hatchery Atlantic salmon smolts. Trans Am Fish Soc. 146(6): 1279-1290.

Lennox RJ, Aarestrup SK, Cooke SJ, Cowley PD, Deng ZD, Fisk AT, Harcourt RG, Heupel M, Hinch SG, Holland KN, Hussey NE, Iverson SJ, Kessel ST, Kocik JF, Lucas MC, Mills Flemming J, Nguyen VM, Stokesbury MJW, Vagle S, Vanderzwaag DL, Whoriskey, FG Young N. 2017. Envisioning the future of aquatic animal tracking: Technology, science, and application. BioScience. 67(10): 884–896.

O’Malley MB, Saunders R, Stevens JR, Jech JM, Sheehan TF. 2017. Using hydroacoustics to describe pelagic fish distribution in the Penobscot Estuary, Maine. Trans Am Fish Soc. 146(5): 817-833.

Goulette G, Hawkes J. 2017. Altering vertical placement of hydroacoustic receivers for improved efficiency in coldwater estuary zones. N Am J Fish Manage. 37(5): 981-988.

Sheehan TF, Davison P, Deschamps D, Drumm A, Millane M, Morgan T, Music P, Niven A, Nygaard R, King TL, Robertson MJ, Ó Maoiléidigh N. 2017. The International Sampling Program: Continent of origin and biological characteristics of Atlantic salmon collected at West Greenland in 2015. 17-13; 27 p.

Renkawitz MD, Trinko Lake TR. 2017. Program and abstracts from the 2016 Atlantic Salmon Ecosystems Forum. 17-04; 59 p.


Lipsky CA, Saunders R, Stevens JR. 2016. Evidence of successful spawning and other life-history aspects of Alosa sapidissima (American shad) in the Penobscot River and Estuary. Northeast Nat. 23(3): 367-377.

Kocik JF. 2016. Book review: Running Silver: Restoring Atlantic Rivers and their Great Fish Migrations. Fisheries. 41(1): 49.

Goulette GS, Lipsky CA. 2016. Nonlethal batch identification of Atlantic salmon using coded wire tags. N Am J Fish Manage. 36(5): 1084-1089.


Renkawitz MD, Sheehan TF, Dixon HJ, Nygaard R. 2015. Changing trophic structure and energy flow in the Northwest Atlantic: implications for Atlantic salmon feeding at West Greenland. Mar Ecol Prog Ser. 538: 197-211.

Bradbury IR, Hamilton LC, Rafferty S, Meerburg D, Poole R, Dempson JB, Robertson MJ, Reddin DG, Bourret V, Dionne M, Chaput G, Sheehan TF, King TL, Berna L. 2015. Genetic mixed-stock analysis disentangles spatial and temporal variation in composition of the West Greenland Atlantic Salmon fishery. ICES J Mar Sci. 73(9): 2311–2321.

Kelly JT, Lerner DT, O’Dea MF, Regish AM, Monette MY, Hawkes JP, Nislow KH, McCormick SD. 2015. Evidence for episodic acidification effects on migrating Atlantic salmon Salmo salar smolts. J Fish Biol. 87(5): 1129-1146.

Stich DS, Kinnison MT, Kocik JF, Zydlewski JD. 2015. Initiation of migration and movement rates of Atlantic salmon smolts in fresh water. Can J Fish Aquat Sci. 72(9): 1339-1351.

Stich DS, Zydlewski GB, Kocik JF, Zydlewski JD. 2015. Linking behavior, physiology, and survival of Atlantic salmon smolts during estuary migration. Mar Coast Fish. 7(1): 68-86.

Renkawitz MD, Trinko Lake TR. 2015. Program and abstracts of the 2014 Atlantic Salmon Ecosystems Forum. 15-11; 57 p.

Hussey NE, Kessel ST, Aarestrup K, Cooke SJ, Cowley PD, Fisk AT, Harcourt RG, Holland KN, Iverson SJ, Kocik JF, Mills Flemming JE, Whoriskey FG. 2015. Aquatic animal telemetry: a window into the underwater world. Science. 348(6240): 1255642.

Sheehan TF, Deschamps D, Trinko Lake T, McKelvey S, Thomas K, Toms S, Nygaard R, King TL, Robertson MJ, Ó Maoiléidigh N. 2015. The International Sampling Program: Continent of origin and biological characteristics of Atlantic salmon collected at West Greenland in 2013. 15-22; 27 p.

Minke-Martin V, Dempson JB, Sheehan TF, Power M. 2015. Otolith-derived estimates of marine temperature use by West Greenland Atlantic salmon (Salmo salar). ICES J Mar Sci. 72(7): 2139-2148.

Sheehan TF, Belleni L, Coghlan B, Deschamps D, Haas-Castro R, Mee D, Thomas K, Nygaard R, King TL, Robertson MJ, Maoiléidigh N. 2015. The International Sampling Program: Continent of origin and biological characteristics of Atlantic salmon collected at West Greenland in 2014. 15-23; 33 p.

Nieland JL, Sheehan TF, Saunders R. 2015. Assessing demographic effects of dams on diadromous fish: a case study for Atlantic salmon in the Penobscot River, Maine. ICES J Mar Sci. 72(8): 2423-2437.


Byron CJ, Pershing AJ, Stockwell JD, Xue H, Kocik, JF. 2014. Migration model of post-smolt Atlantic salmon (Salmo salar) in the Gulf of Maine. Fish Oceanogr. 23: 172-189.

Good MJ, Kocik JF. 2014. The intern-mentor experience: a sample of the real world. In: Taylor WW, Lynch AJ, Léonard NJ, editors. Future of Fisheries: Perspectives for Emerging Professionals. Bethesda (MD): Am Fish Soc p 31-38.

Goulette G, Hawkes J, Kocik J, Manning J, Music P, Wallinga J, Zydlewski G. 2014. Opportunistic acoustic telemetry platforms: Benefits of collaboration in the Gulf of Maine. Fisheries. 39(10): 441-450.

Ardren WR, Busack C, Kocik JF. 2014. Collaboration between Atlantic and Pacific salmon biologists to enhance recovery of endangered salmon in North America. Rev Fish Biol Fish. 24: 685-688.

Hayes S, Kocik J. 2014. Comparative estuarine and marine migration ecology of Atlantic salmon and steelhead: blue highways and open plains. Rev Fish Biol Fish. 24: 757-780.

Miller AS, Miller TJ, Mills KE, Sheehan TF. 2014. Retrospective analysis of Atlantic salmon (Salmo salar) marine growth and condition in the northwest Atlantic based on tag-recovery data. Fish Oceanogr. 23(2): 103-115.

Otero J, L'Abée-Lund JH, Castro-Santos T, Leonardsson K, Storvik GO, Jonsson B, Dempson B, Russell IC, Jensen AJ, Baglinière J-L, Dionne M, Armstrong JD, Romakkaniemi A, Letcher BH, Kocik JF, Erkinaro J, Poole R, Rogan G, Lundqvist H, MacLean JC, Jokikokko E, Arnekleiv JV, Kennedy RJ, Niemelä E, Caballero P, Music PA, Antonsson T, Gudjonsson S, Veselov AE, Lamberg A, Groom S, Taylor BH, Taberner M, Dillane M, Arnason F, Horton G, Hvidsten NA, Jonsson IR, Jonsson N, McKelvey S, Naesje TF, Skaala O, Smith GW, Saegrov H, Stenseth NC, Vøllestad LA. 2014. Basin-scale phenology and effects of climate variability on global timing of initial seaward migration of Atlantic salmon (Salmo salar). Global Change Biol. 20: 61-75.

Moustahfid H, Weise M, Simmons S, Block B, Holland K, Ault J, Kocik J, Costa D, Lindley ST, Mate B, Hayes S, Holbrook CM, Seitz A, Arendt M, Payne J, Mahmoudi B, Alexander C, Moore P, Price J, Wilson D. 2014. Meeting our Nation’s needs for biological and environmental monitoring: strategic plan and recommendations for a national telemetry network (ATN) through U.S. IOOS. 534; 7 p.

Weitkamp L, Goulette G, Hawkes J, O'Malley M, Lipsky C. 2014. Juvenile salmon in estuaries: comparisons between Atlantic and Pacific salmon. Rev Fish Biol Fish. 24(3): 713-736.


McCormick SD, Sheehan T, Björnsson BT, Lipsky C, Kocik J, Regish AM, O’Dea MF. 2013. Physiological and endocrine changes in Atlantic salmon smolts during hatchery rearing, downstream migration, and ocean entry. Can J Fish Aquat Sci. 70: 105-118.

Kocik JF, Lipsky C, Miller T, Rago P, Shepherd G. 2013. An Atlantic sturgeon population index for ESA management analysis. 13-06; 36 p.

Hawkes JP, Saunders R, Vashon A, Cooperman MS. 2013. Assessing efficacy of non-lethal harassment of double-crested cormorants to improve Atlantic salmon smolt survival. Northeast Nat. 20(1): 1-18.

Nieland JL, Sheehan TF, Saunders R, Murphy JS, Trinko Lake TR, Stevens JR. 2013. Dam impact analysis model for Atlantic salmon in the Penobscot River, Maine. 13-09; 524 p.

Sheehan TF, Assunção MGL, Deschamps D, Laughton B, Ó Cuaig M, Nygaard R, King TL, Robertson MJ, Ó Maoiléidigh N. 2013. The International Sampling Program: Continent of origin and biological characteristics of Atlantic salmon collected at West Greenland in 2012. 13-20; 25 p.


Lacroix GL, Knox D, Sheehan TF, Renkawitz MD, Bartron ML. 2012. Distribution of US Atlantic salmon postsmolts in the Gulf of Maine. Trans Am Fish Soc. 141: 934-942.

Lipsky C, Hastings E, Brown R, Dill R. 2012. Correcting visual implant elastomer tag identification to better evaluate adult Atlantic salmon returns from multiple stocking sites in the Penobscot River, Maine, USA. In: McKenzie J, Parsons B, Seitz A, Kopf RK, Mesa M, Phelps Q, editors. Advances in Fish Tagging and Marking Technology Am Fish Soc Symp. 76: 63-75.

Miller AS, Sheehan TF, Renkawitz MD, Meister AL, Miller TJ. 2012. Revisiting the marine migration of US Atlantic salmon using historical Carlin tag data. ICES J Mar Sci. 69: 1609-1615.

Reddin DG, Hansen LP, Bakkestuen V, Russell I, White J, Potter ECE, Sheehan T, Ó Maoiléidigh N, Dempson JB, Smith GW, Isaksson A, Fowler M, Jacobsen JA, Mork KA, Amiro P. 2012. Distribution and biological characteristics of Atlantic salmon (Salmo salar) at Greenland based on the analysis of historical tag recoveries. ICES J Mar Sci. 69(9): 1589-1597.

MacLean SA. 2012. Programs and abstracts of the Maine Atlantic Salmon Forums. 12-12; 171 p.

Renkawitz MD, Sheehan TF, Goulette GS. 2012. Swimming depth, behavior, and survival of Atlantic salmon postsmolts in Penobscot Bay, Maine. Trans Am Fish Soc. 141(5): 1219-1229.

Miller AS, Sheehan TF, Spencer RC, Renkawitz MD, Friedland KD, Meister AL. 2012. Description of the historic US Atlantic salmon (Salmo salar L.) tagging programs and subsequent databases. 12-13; 49 p.

Caldarone EM, MacLean SA, Sharack B. 2012. Evaluation of bioelectrical impedance analysis and Fulton’s condition factor as nonlethal techniques for estimating short-term responses in postsmolt Atlantic salmon (Salmo salar) to food availability. Fish Bull. 110: 257-270.

Sheehan TF, Reddin DG, Chaput G, Renkawitz MD. 2012. SALSEA North America: a pelagic ecosystem survey targeting Atlantic salmon in the Northwest Atlantic. ICES J Mar Sci. 69(9): 1580-1588.

Sheehan TF, Assunção MGL, Chisholm N, Deschamps D, Dixon H, Renkawitz M, Rogan G, Nygaard R, King TL, Robertson MJ, O' Maoiléidigh N. 2012. The International Sampling Program, continent of origin and biological characteristics of Atlantic salmon (Salmo salar) collected at West Greenland in 2011. 12-24; 27 p.

Dixon H, Power M, Dempson JB, Sheehan TF, Chaput G. 2012. Characterizing the trophic position and shift in Atlantic salmon (Salmo salar) from freshwater to marine life-cycle phases using stable isotopes. ICES J Mar Sci. 69(9): 1646-1655.


Moustahfid H, Grimes C, Kocik J, Block B, Holland K, Payne J, Fox D, Seitz A, Alexander C. 2011. Toward a National Animal Telemetry Observing Network (ATN) for our oceans, coasts and great lakes: workshop synthesis report. U.S. IOOS workshop, March 2011, Santa Cruz, California 482; 51 p.

Sheehan TF, Renkawitz MD, Brown RW. 2011. Surface trawl survey for U.S. origin Atlantic salmon Salmo salar. J Fish Biol. 79: 374-398.

O’Dor R, Acosta J, Bergstad OA, Brainard R, Brattey J, Canals M, Costa D, Gunn J, Horne JK, Iken K, Kocik J, Konar B, Payne J, Reid C, Robison B, Steinke D, Vanden Berg E. 2011. Bringing life to ocean observation. In: Hall J, Harrison DE, Stammer D, eds. Proceedings of OceanObs’09: Sustained Ocean Observations and Information for Society (Vol. 1), Venice, Italy, 21-25 September 2009 306.

Renkawitz MD, Sheehan TF. 2011. Feeding ecology of early marine phase Atlantic salmon Salmo salar post-smolts. J Fish Biol. 79: 356-373.

Renkawitz MD, Gregory RS, Schneider DC. 2011. Habitat dependent growth of three species of bottom settling fish in a coastal fjord. J Exp Mar Biol Ecol. 409: 79-88.


Music PA, Hawkes JP, Cooperman MS. 2010. Magnitude and causes of smolt mortality in rotary screw traps: an Atlantic salmon case study. N Am J Fish Manage. 30: 713-722.


Sheehan TF, Legault CM, King TL, Spidle AP. 2009. Probabilistic-based genetic assignment model: assignments to subcontinent of origin of the West Greenland Atlantic salmon harvest. ICES J Mar Sci. 67(3): 537-550.

Kocik JF, Hawkes JP, Sheehan TF, Music PA, Beland KF. 2009. Assessing estuarine and coastal migration and survival of wild Atlantic salmon smolts from the Narraguagus River, Maine, using ultrasonic telemetry. In: Haro AJ, Smith KL, Rulifson RA, Moffitt CM, Klauda RJ, Dadswell MF, Cunjak RA, Cooper JE, Beal KL, Aver TS, editors. Challenges for Diadromous Fishes in a Dynamic Global Environment. Am Fish Soc Symp. 69: 293-310.


Colligan M, Sheehan T, Pruden J, Kocik JF. 2008. The challenges posed by international management of Atlantic salmon: balancing commercial, recreational and societal interests. In: Schechter MG, Leonard NJ, Taylor WW, eds. International Governance of Fisheries Ecosystems: Learning from the Past, Finding Solutions for the Future. Bethesda [MD]; American Fisheries Society Chapter 15.


MacLean SA. 2006. Salmonid pathogens in northwestern Atlantic fishes. 2006-001; 7 p.

Friedland KD, Clarke LM, Dutil J-D, Salminen M. 2006. The relationship between smolt and postsmolt growth for Atlantic salmon (Salmo salar) in the Gulf of St. Lawrence [Note]. Fish Bull. 104(1): 149-155.

Saunders R, Hachey MA, Fay CW. 2006. Maine's Diadromous Fish Community: Past, Present, and Implications for Atlantic Salmon Recovery. Fisheries. 31(11): 537-547.[537:MDFC]2.0.CO;2

Haas-Castro RE, Sheehan TF, Cadrin SX, Trial JG. 2006. Scale pattern analysis discriminates Atlantic salmon by river reach rearing origin. N Am J Fish Manage. 26: 289-300.

Reddin DG, Friedland KD, Downton P. 2006. Early marine use of thermal habitat by Atlantic salmon smolts (Salmo salar L.). Fish Bull. 104: 415-428.

Sweka JA, Legault CM, Beland KF, Trial J, Millard MJ. 2006. Evaluation of removal sampling for basinwide assessment of Atlantic salmon. N Am J Fish Manage. 26: 995-1002.


Sheehan TF, Kocik JF, Cadrin SX, Legault CM, Atkinson E, Bengtson D. 2005. Marine growth and morphometrics for three populations of Atlantic salmon from Eastern Maine, USA. Trans Am Fish Soc. 134: 775-788.

Chaput G, Legault CM, Reddin DG, Caron F, Amiro PG. 2005. Provision of catch advice taking account of non-stationarity in productivity of Atlantic salmon (Salmo salar L.) in the northwest Atlantic. ICES J Mar Sci. 62: 131-143.

Friedland KD, Chaput G, MacLean JC. 2005. The emerging role of climate in post-smolt growth of Atlantic salmon. ICES J Mar Sci. 62: 1338-1349.

Giray C, Opitz HM, MacLean SA, Bouchard D. 2005. Comparison of lethal versus non-lethal sample sources for the detection of infectious salmon anemia virus (ISAV). Dis Aquat Org. 66: 181-185.

Nérette P, Dohoo I, Hammell L, Gagné N, Barbash P, MacLean S, Yason C. 2005. Estimation of the repeatablility and reproducibility of three diagnostic tests for infectious Salmon anaemia virus. J Fish Dis. 28: 101-110.


Beland KF, Trial JG, Kocik JF. 2004. Use of riffle and run habitats with aquatic vegetation by juvenile Atlantic salmon. N Am J Fish Manage. 24: 525-533.

Clarke LM, Friedland KD. 2004. Influence of growth and temperature on strontium deposition in the otoliths of Altantic salmon, Salmo salar L. J Fish Biol. 65: 744-759.

FitzGerald JL, Sheehan TF, Kocik JF. 2004. Visibility of visual implant elastomer tags in Atlantic salmon reared for two years in marine net-pens. N Am J Fish Manage. 24(1): 222-227.

Kocik JF. 2004. North Atlantic climate book: a one-stop reference for a complicated subject. Trends Ecol Evol. 19(12): 624-625.

Reddin DG, Friedland KD, Downton P, Dempson JB, Mullins CC. 2004. Thermal habitat experienced by Atlantic salmon (Salmo salar L.) kelts in coastal Newfoundland waters. Fish Oceanogr. 13: 24-35.


Magee JA, Obedzinski M, McCormick SD, Kocik JF. 2003. Effects of episodic acidification on Atlantic salmon (Salmo salar) smolts. Can J Fish Aquat Sci. 60: 214-221.

Spidle AP, Kalinowski ST, Lubinski BA, Perkins DL, Beland KF, Kocik JF, King TL. 2003. Population structure of Atlantic salmon in Maine with reference to populations from Atlantic Canada. Trans Am Fish Soc. 132: 196-209.

Wells BK, Friedland KD, Clarke LM. 2003. Increment patterns in otoliths and scales from mature Atlantic salmon Salmo salar. Mar Ecol Prog Ser. 262: 293-298.

MacLean SA, Bouchard DA, Ellis SK. 2003. Survey of nonsalmonid marine fishes for detection of infectious salmon anemia virus and other salmon pathogens. In: Miller O; Rocco C, tech coords. International response to infectious salmon anemia: prevention, control, and eradication. Proceedings of a symposium; 2002 Sep 3-4; New Orleans LA; p 135-143.

Friedland KD, Reddin DG, Castonguay M. 2003. Ocean thermal conditions in the post-smolt nursery of North American Atlantic salmon. ICES J Mar Sci. 60(2): 343-355.

Friedland KD, Reddin DG, McMenemy JR, Drinkwater KF. 2003. Multidecadal trends in North American Atlantic salmon (Salmo salar) stocks and climate trends relevant to juvenile survival. Can J Fish Aquat Sci. 60: 563-583.


Kocik JF, Friedland KD. 2002. Salmons and trouts, order Salminiformes. In: Collette, BB; Klein-MacPhee, G, eds  Bigelow and Schroeder's Fishes of the Gulf of Maine  3rd ed  Washington, DC: Smithsonian Institution Press; p 170-181.

Orrell TM, Carpenter KE, Musick JA, Graves JE. 2002. Phylogenetic and biogeographic analysis of the Sparidae (Perciformes: Percoidei) from Cytochrome b sequences. Copeia. 2002(3): 618-631.

Friedland KD. 2002. Emerging science and management issues with Atlantic salmon. In: Lynch, KD; Jones, ML; Taylor, WW, eds. Sustaining North American Salmon: Perspective Across Regimes and Disciplines Bethesda, MD: American Fisheries Society p 117-136.

Kocik JF, Brown RW. 2002. From game fish to tame fish: Atlantic salmon in North America, 1798 to 1998. In: Lynch, KD; Jones, ML; Taylor, WW Sustaining North American salmon: perspectives across regions and disciplines Bethesda, MD: American Fisheries Society; p 3-31.


Beland KF, Kocik JF, vandeSande J, Sheehan TF. 2001. Striped bass predation upon Atlantic salmon smolts in Maine. Northeast Nat. 8(3): 267-274.

Friedland KD, Walker RV, Davis ND, Myers KW, Boehlert GW, Urawa S, Ueno Y. 2001. Open-ocean orientation and return migration routes of chum salmon based on temperature data from storage tags. Mar Ecol Prog Ser. 216: 235-252.

Magee JA, Haines TA, Kocik JF, Beland KF. 2001. Effects of acidity and aluminum on the physiology and migratory behavior of Atlantic salmon smolts in Maine, USA. Water Air Soil Pollut. 130: 881-886.


Friedland KD, Reddin DG. 2000. Growth patterns of Labrador Sea Atlantic salmon postsmolts and the temporal scale of recruitment synchrony for North American salmon stocks. Can J Fish Aquat Sci. 57: 1181-1189.

Walker RV, Myers KW, Davis ND, Aydin KY, Friedland KD, Carlson HR, Boehlert GW, Urawa S, Ueno Y, Anma G. 2000. Diurnal variation in thermal environment experienced by salmonids in the North Pacific as indicated by data storage tags. Fish Oceanogr. 9: 171-186.

Friedland KD, Hansen LP, Dunkley DA, MacLean JC. 2000. Linkage between ocean climate, post-smolt growth, and survival of Atlantic salmon (Salmo salar L.) in the North Sea area. ICES J Mar Sci. 57: 419-429.


Friedland KD, Dutil J-D, Sadusky T. 1999. Growth patterns in postsmolts and the nature of the marine juvenile nursery for Atlantic salmon, Salmo salar. Fish Bull. 97: 472-481.

Jonsson B, Waples RS, Friedland KD. 1999. Extinction considerations for diadromous fishes. ICES J Mar Sci. 56: 405-409.

Kocik JF, Jones ML. 1999. Pacific salmonines in the Great Lakes Basin. In: Taylor WW, Ferreri CP, editors. Great Lakes fisheries policy and management; a binational perspective. Michigan State University Press


Kocik JF, Ferreri CP. 1998. Juvenile production variation in salmonids: population dynamics, habitat, and the role of spatial relationships. Can J Fish Aquat Sci. 55(Suppl 1): 191-200.

Sheehan TF. 1998. First record of the ragged-tooth shark, Odontaspis ferox, off the U.S. Atlantic coast. Mar Fish Rev. 60(1): 33-34.

Dodson JJ, Gibson R, Cunjak R, Friedland KD, De Leaniz C, Gross M, Newbury R, Nielsen J, Power M, Roy S. 1998. Elements in the development of conservation plans for Atlantic salmon (Salmo salar). Can J Fish Aquat Sci. 55(S1): 312-323.

Blackwell BF, Gries G, Juanes F, Friedland KD, Stolte LW, McKeon JF. 1998. Simulating migration mortality of Atlantic salmon smolts in the Merrimack River. N Am J Fish Manage. 18: 31-45.

Friedland KD, Hansen LP, Dunkley DA. 1998. Marine temperatures experienced by postsmolts and the survival of Atlantic salmon, Salmo salar L., in the North Sea area. Fish Oceanogr. 7(1): 22-34.

Friedland KD, Reddin DG, Shimizu N, Haas RE, Youngson AF. 1998. Strontium:calcium ratios in Atlantic salmon (Salmo salar) otoliths and observations on growth and maturation. Can J Fish Aquat Sci. 55: 1158-1168.

Friedland KD. 1998. Ocean climate influences on critical Atlantic salmon (Salmo salar) life history events. Can J Fish Aquat Sci. 55(S1): 119-130.


Friedland KD, Haas RE. 1996. Marine post-smolt growth and age at maturity of Atlantic salmon. J Fish Biol. 48: 1-15.

Friedland KD, Haas RE, Sheehan TF. 1996. Post-smolt growth, maturation, and survival of two stocks of Atlantic salmon. Fish Bull. 94(4): 654-663.


Kocik JF, Taylor WW. 1994. Summer survival and growth of brown trout with and without steelhead under equal total salmonine densities in an artificial stream. Trans Am Fish Soc. 123: 931-938.

Friedland KD, Esteves C, Hansen LP, Lund RA. 1994. Discrimination of Norwegian farm, ranch, and wild-origin Atlantic salmon Salmo salar L. by image processing. Fish Manage Ecol. 1: 117-128.

Friedland KD, Reddin DG. 1994. The use of otolith morphology in stock discriminations of Atlantic salmon Salmo salar. Can J Fish Aquat Sci. 51: 91-98.


Friedland KD, Reddin DG, Kocik JF. 1993. Marine survival of North American and European Atlantic salmon: effects of growth and environment. ICES J Mar Sci. 50: 481-492.


Bermingham E, Forbes SH, Friedland KD, Pla C. 1991. Discrimination between Atlantic salmon (Salmo salar) of North American and European origin using restriction analyses of mitochondrial DNA. Can J Fish Aquat Sci. 48(5): 884-893.